]> GH barcode-00005499 Costa Rica Guanacaste Lower slopes of Cerro Santamaría, north of farmhouse, Hacienda Santamaría; 720-920 m. Piper papulaecaule Trelease ex Standley Trelease ex Standley 2010-08-16 16:43:04 C. W. Dodge & W. S. Thomas